Adult body sizes (not including the horn) vary between 50 and 85 mm (2. 0 and 3. 3 in) in length and 29 and 42 mm (1. 1 and 1. 7 in) in width, [clarification needed] though male Hercules beetles may reach up to 175 mm (7 in) in length (including the horn), making them the longest species of beetle in the world, if jaws and/or horns are included in the measurement. The size of this horn is naturally very variable; more so than any variation of the size of legs, wings, or overall body size in the species. This variability results from developmental mechanisms that couple genetic predisposition with nutrition, stress, exposure to parasites, and/or physiological conditions.
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